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Dr. Phil Trathan - Profile


158 peer-reviewed publications (papers and book chapters), with 2 more in press; H-index 33.

In Press

Hart T., Trathan P.N., Brown R.M., Teacher A.G.F., Clucas G.V., Ahmed S., Rogers, A.D. (in press). Gene flow in a philopatric seabird; macaroni penguin populations around South Georgia. Polar Biology.
Ratcliffe N., Trathan P.N. (in press). A review of the diet and foraging movements of penguins breeding within the CCAMLR area. CCAMLR Science.
Southwell1 C., Bengtson J., Bester M., Blix A.S., Bornemann H., Boveng P., Cameron M., Forcada J., Laake J., Nordøy E., Plötz J., Rogers T., Southwell D., Steinhage D., Stewart B.S., Trathan P.N. (in press). A review of data on abundance, trends in abundance, habitat use and diet of ice-breeding seals in the Southern Ocean. CCAMLR Science.

Fretwell, P.T., Trathan, P.N., Wienecke, B. & Kooyman, G.L. (2014). Emperor penguins breeding on iceshelves. PLOS ONE 9(1): e85285. doi:10.1371/journal.pone.0085285
Larsen, J.N., Anisimov, O.A., Constable, A., Hollowed, A., Maynard, N., Prestrud, P., Prowse, T., Stone, J., Callaghan, T., Carey, M., Convey, P., Derocher, A., Fretwell, P.T., Forbes, B.C., Glomsrød, S., Hodgson, D., Hofmann, E., Hovelsrud, G.K., Ljubicic, G.L., Loeng, H., Murphy, E., Nicol, S., Oskal, A., Reist, J.D., Trathan, P.N., Weinecke, B. & Wrona, F. (2014). Chapter 28. Polar Regions. In: Working Group II Climate Change 2014: Impacts, Adaptation, and Vulnerability. IPCC AR5.
Thiebot, J.B., Authier, M., Trathan, P.N. & Bost, C.A. (2014). Gentlemen first? 'Broken stick' modelling reveals sex-related homing decision date in migrating seabirds. Journal of Zoology 292 (1): 25-30.

Crossin G., Trathan P.N., Crawford R. (2013). The macaroni and royal penguin. In: Biology and Conservation of the World’s Penguins (eds. P.D. Boersma and P.G. Borboroglu). University of Washington Press, Seattle.
Dewar, M.L., Arnould, J.P.Y., Dann, P., Trathan, P.N., Groscolas, R. & Smith, S. (2013). Interspecific variations in the gastrointestinal microbiota in penguins. MicrobiologyOpen 2(1): 195-204.
Grant, S.M., Hill, S.L., Trathan, P.N. & Murphy, E.J. (2013). Ecosystem services of the Southern Ocean: trade-offs in decision-making. Antarctic Science 25(5): 603-617.
Hoffman J.I., Thorne M.A.S., Trathan P.N., Forcada J. (2013). Transcriptome of the dead: characterisation of immune genes and marker development from necropsy samples in a free-ranging marine mammal. BMC Genomics 14: 52. doi: 10.1186/1471-2164-14-52.
Kazama, K., Hirata, K., Yamamoto, T., Hashimoto, H., Takahashi, A., Niizuma, Y., Trathan, P.N. & Watanuki, Y. (2013). Movements and activities of male black-tailed gulls in breeding and sabbatical years. Journal of Avian Biology 44(6): 603-608.
Lynch H.J., Ratcliffe N., Passmore J., Foster E., Trathan P.N. (2013). Sensitivity analysis identifies high influence sites for estimates of penguin krill consumption on the Antarctic Peninsula. Antarctic Science 25: 19-23.
Murphy, E.J., Hofmann, E.E., Watkins, J.L., Johnston, N.M., Pinones, A., Ballerini, T., Hill, S.L., Trathan, P.N., Tarling, G.A., Cavanagh, R.A., Young, E.F., Thorpe, S.E. & Fretwell, P. (2013). Comparison of the structure and function of Southern Ocean regional ecosystems: The Antarctic Peninsula and South Georgia. Journal of Marine Systems 109(SI): 22-42.
Ratcliffe, N., Takahashi, A., O'Sullivan, C., Adlard, S., Trathan, P.N., Harris, M.P. & Wanless, S. (2013). The roles of sex, mass and individual specialisation in partitioning foraging-depth niches of a pursuit-diving predator. PLOS ONE 8(10): e79107. doi: 10.1371/journal.pone.0079107.
Sakamoto, K.Q., Takahashi, A., Iwata, T., Yamamoto, T., Yamamoto, M. & Trathan, P.N. (2013). Heart rate and estimated energy expenditure of flapping and gliding in black-browed albatrosses. Journal of Experimental Biology 216(16): 3175-3182.
Schmale, J., Schneider, J., Nemitz, E., Tang, Y.S., Dragosits, U., Blackall, T.D., Trathan, P.N., Phillips, G.J., Sutton, M. & Braban, C.F. (2013). Sub-Antarctic marine aerosol: dominant contributions from biogenic sources. Atmospheric chemistry and physics 13(17): 8669-8694.
Thiebot, J.B., Cherel, Y., Crawford, R.J.M., Makhado, A.B., Trathan, P.N., Pinaud, D. & Bost, C.-A. (2013). A space oddity: Geographic and specific modulation of migration in Eudyptes penguins. PLOS ONE 8(8): e71429. doi: 10.1371/journal.pone.0071429.
Trathan P.N., Ballard G. (2013). The Adélie penguin. In: Biology and Conservation of the World’s Penguins (eds. P.D. Boersma and P.G. Borboroglu). University of Washington Press, Seattle.
Trathan P.N., Grant S.M., Siegel V., Kock K.-H. (2013). Precautionary spatial protection to facilitate the scientific study of habitats and communities under ice shelves in the context of recent, rapid, regional climate change. CCAMLR Science 20: 139-151.


Crossin G.T., Dawson A., Phillips R.A., Trathan P.N., Gorman K.B., Adlard S., Williams T.D. (2012). Seasonal patterns of prolactin and corticosterone secretion in an Antarctic seabird that moults during reproduction. General and Comparative Endocrinology 175: 74-81.
Crossin G.T., Phillips R.A., Trathan P.N., Fox D.S., Dawson A., Wynne-Edwards K.E., Williams T.D. (2012). Migratory carryover effects and endocrinological correlates of reproductive decisions and reproductive success in female albatrosses. General and Comparative Endocrinology 176: 151-157.
Crossin G.T., Trathan P.N., Phillips R.A., Gorman K.B., Dawson A., Sakamoto K.Q., Williams T.D. (2012). Corticosterone predicts foraging behavior and parental care in macaroni penguins. American Naturalist 180: E31-E41.
Crossin G., Trathan P.N., Williams, T.D. (2012). Potential mode of clutch size determination and follicle development in Eudyptes penguins. Polar Biology 35: 313-317.
Flores H., Atkinson A., Kawaguchi S., Krafft B.A., Milinevsky G., Nicol S., Reiss C., Tarling G.A., Werner R., Rebolledo E.B., Cirelli V., Cuzin-Roudy J., Fielding S., Groeneveld J.J., Haraldsson M., Lombana A., Marschoff E., Meyer B., Pakhomov E.A., Rombola E., Schmidt K., Siegel V., Teschke M., Tonkes H., Toullec J.Y., Trathan P.N., Tremblay N., Van de Putte A.P., van Franeker J.A., Werner T. (2012). Impact of climate change on Antarctic krill. Marine Ecology-Progress Series 458: 1-19.
Forcada J., Trathan P.N., Boveng P.L., Boyd I.L., Burns J.M., Costa D.P., Fedak M., Rogers T.L., Southwell C.J. (2012). Responses of Antarctic pack-ice seals to environmental change and increasing krill fishing. Biological Conservation 149: 40-50.
Fretwell P.T., LaRue M.A. Morin P., Kooyman G.L., Wienecke B., Ratcliffe N., Fox A.J., Fleming A.H., Porter C., Trathan P.N. (2012). An emperor penguin population estimate: The first global, synoptic survey of a species from space. PLoS ONE 7: e33751. doi: 10.1371/journal.pone.0033751.
Grant S.M., Convey P., Hughes K.A., Phillips R.A., Trathan P.N. (2012). Conservation of Antarctic marine and terrestrial ecosystems. In An Extreme Environment in a Changing World (eds. A. Rogers, N. Johnston, A. Clarke and E.J. Murphy). Blackwell Publishing, pp 492-525.
Iwata T., Sakamoto K.Q., Takahashi A., Edwards E.W.J., Staniland I.J., Trathan P.N., Naito N. (2012). Using a mandible accelerometer to study fine-scale foraging behavior of free-ranging Antarctic fur seals. Marine Mammal Science 28: 345-357.
Lynch H.J., Naveen R., Trathan P.N., Fagan W.F. (2012). Spatially integrated assessment reveals widespread changes in penguin populations on the Antarctic Peninsula. Ecology 93:1367-1377.
Murphy E.J., Cavanagh R.D., Hofmann E.E., Hill S.L., Constable A.J., Costa D.P., Pinkerton M.H., Johnston N.M., Trathan P.N., Klinck J.M., Wolf-Gladrow D.A., Daly K.L., Maury O., Doney S.C. (2012) Developing integrated models of Southern Ocean food webs: Including ecological complexity, accounting for uncertainty and the importance of scale. Progress in Oceanography 102: 74-92.
Murphy E.J., Hofmann E.E., Watkins J.L., Johnston N.M., Pinones A., Ballerini T., Hill S.L., Trathan P.N., Tarling G.A., Cavanagh R.A., Young E.F., Thorpe S.E., Fretwell P. (2013). Comparison of the structure and function of Southern Ocean regional ecosystems: The Antarctic Peninsula and South Georgia. Journal of Marine Systems 109: 22- 42.
Murphy E.J., Watkins J.L., Trathan P.N., Reid K., Meredith M.P., Hill S.L., Thorpe S.E., Johnston N.M., Clarke A., Tarling G.A., Collins M.A., Forcada J., Atkinson A., Ward P., Staniland I.J., Pond D.W., Cavanagh R.A., Shreeve R.S., Korb R.E., Whitehouse M.J. , Rodhouse P.G., Enderlein P., Hirst A.G., Martin A.R., Briggs D.R., Cunningham N.J., Fleming A.H. (2012). Spatial and Temporal Operation of the Scotia Sea Ecosystem. In An Extreme Environment in a Changing World (eds. A. Rogers, N. Johnston, A. Clarke and E.J. Murphy). Blackwell Publishing, pp 160-212.
Shirai M., Yamamoto M., Ebine N., Yamamoto T., Trathan P.N., Yoda K., Oka N., Niizuma Y. (2012). Basal and field metabolic rates of Streaked Shearwater during the chick-rearing period. Ornithological Science 11: 47- 55.
Staniland I.J., Robinson S.L., Silk J.R.D., Warren N., Trathan P.N. (2012). Winter distribution and haul-out behaviour of female Antarctic fur seals from South Georgia. Marine Biology 159: 291-301.
Thiebot J.B., Cherel Y., Trathan, P.N., Bost C.A. (2012). Coexistence of oceanic predators on wintering areas explained by population-scale foraging segregation in space or time. Ecology 93: 122-130.
Trathan P.N. (2012). Settle discord over the Southern Ocean. Nature 492: 186-186.
Trathan P.N. (2012). Breeding penguins and Vagrant penguins in Burton R. (Ed.) A Field Guide to the Wildlife of South Georgia. South Georgia Heritage Trust.
Trathan P.N., Forcada J., Murphy E.J. (2012). Environmental forcing and Southern Ocean marine predator populations: effects of climate change and variability. In: An Extreme Environment in a Changing World (eds. A. Rogers, N. Johnston, A. Clarke and E.J. Murphy). Blackwell Publishing, pp 335-353.
Trathan P.N., Ratcliffe N., Masden E.A. (2012). Ecological drivers of change at South Georgia: the krill surplus, or climate variability. Ecography 35: 983-993.
Yamamoto T., Takahashi A., Oka N., Shirai, M., Yamamoto M., Katsumata N., Sato K., Watanabe S., Trathan P.N. (2012). Inter-colony differences in the incubation pattern of streaked shearwaters in relation to the local marine environment Waterbirds 35: 248-259.

Dunn M.J., Silk J.R.D., Trathan P.N. (2011). Post-breeding dispersal of Adélie penguins (Pygoscelis adeliae) nesting at Signy Island, South Orkney Islands. Polar Biology 34: 205-214. doi: 10.1007/s00300-010-0870-4.
Thiebot J.-B., Cherel Y., Trathan P.N., Bost C.-A. (2011). Inter-population segregation in the wintering areas of macaroni penguins. Marine Ecology-Progress Series 421: 279-290.
Thiebot J.-B., Lescroël A., Pinaud D., Trathan P.N., Bost C.-A. (2011). Larger foraging range but similar habitat selection in non-breeding versus breeding sub-Antarctic penguins. Antarctic Science 23: 117-126.
Trathan P.N., Fretwell, P.T., Stonehouse, B (2011). First recorded loss of an emperor penguin colony in the recent period of Antarctic regional warming: implications for other colonies. PLoS ONE 6: e14738.
Wakefield E.D, Phillips R.A., Trathan P.N., Arata, J. Gales R., Huin N., Robertson G., Waugh S.M., Weimerskirch H., Matthiopoulos J. (2011). Habitat preference, accessibility and competition limit the global distribution of breeding black-browed albatrosses. Ecological Monographs 81: 141-167. doi: 10.1890/09-0763.1.
Yamamoto T., Takahashi A., Oka N., Iida T., Katsumata N., Sato K., Trathan P.N. (2011). Foraging areas of streaked shearwaters in relation to seasonal changes in the marine environment of Northwestern Pacific: Inter-colony and sexual differences. Marine Ecology-Progress Series 424: 191-204.

Ballard G., Toniolo V., Ainley D.G., Parkinson C.L., Arrigo K.R., Trathan P.N. (2010). Responding to climate change: Adélie Penguins confront astronomical and ocean boundaries. Ecology 91: 2056-2069.
Crossin G.T., Trathan P.N., Phillips R.A., Dawson A., Le Bouard F., Williams T.D. (2010). A carryover effect of migration underlies individual variation in reproductive readiness and extreme egg size dimorphism in macaroni penguins. American Naturalist 176: 357-66. 26.
Crossin G.T., Trathan P.N., Phillips R.A., Williams T.D. (2010). Trade-off between migration and reproduction, and the physiological basis of egg size dimorphism in macaroni penguins. Integrative and Comparative Biology 50: Supplement 1, E36-E36.
Hart T., Coulson T., Trathan P.N. (2010). Time series analysis of biologging data: autocorrelation reveals periodicity of diving behaviour in macaroni penguins. Animal Behaviour 79: 845-855.
Hart T., Mann R., Coulson T., Pettorelli N., Trathan P.N. (2010) Behavioural switching in a central place forager: patterns of diving behaviour in the macaroni penguin (Eudyptes chrysolophus). Marine Biology 157: 1543-1553. 31.
Hill S.L., Forcada J., Trathan P.N., Waluda C.M. (2010). Using ecosystem monitoring data to detect impacts. CCAMLR Science 17: 197-212.
Hoffman J.I., Hanson N., Forcada J., Trathan P.N., Amos W. (2010). Getting Long in the Tooth: A Strong Positive Correlation between Canine Size and Heterozygosity in Antarctic Fur Seals Arctocephalus gazelle. Heredity 101: 527-538.
Mori Y., Takahashi A., Trathan P.N., Watanuki Y. (2010). Optimal stroke frequency during diving activity in seabirds. Aquatic Biology 8: 247-257
Reid, K., Watkins, J.L., Murphy, E.J., Trathan, P.N., Fielding, S., Enderlein, P. (2010). Krill population dynamics at South Georgia: implications for ecosystem-based fisheries management. Marine Ecology-Progress Series 399: 243-252.
Scheffer A., Trathan P.N., Collins M. (2010). Foraging behaviour of King Penguins (Aptenodytes patagonicus) in relation to predictable mesoscale oceanographic features in the Polar Front Zone to the north of South Georgia. Progress in Oceanography 86: Special Issue: Sp. Iss. SI 232-245.
Trathan P.N., Agnew D. (2010).Climate change and the Antarctic marine ecosystem: an essay on management implications. Antarctic Science 22: 387-398.
Waluda, C.M., Collins, M.A., Black, A.D., Staniland, I.J., Trathan, P.N. (2010). Linking predator and prey behaviour: contrasts between Antarctic fur seals and macaroni penguins at South Georgia. Marine Biology 157: 99- 112. 10.1007/s00227-009-1299-6.
Watanuki Y., Takahashi A., Trathan P.N., Wanless S., Sakamoto K.Q., Sato K. (2010). Regulation of wing or foot strokes in deep-diving seabirds: a comparison between South Georgian Shag, Common Murre and Macaroni Penguin. Japanese Journal of Ornithology, 59: 20-30. 49.
Yamamoto T., Takahashi A, Katsumata N., Sato K., Trathan P.N. (2010). At-sea distribution and behavior of streaked shearwaters (Calonectris leucomelas) during non-breeding period: from temperate to tropical oceans. Auk, 127: 871-881.

Ahmed, S., Hart, T., Dawson, D.A., Horsburgh, G.J., Trathan, P.N., Rogers, A.D. (2009). Isolation and characterization of macaroni penguin (Eudyptes chrysolophus) microsatellite loci and their utility in other penguin species (Spheniscidae, Aves). Molecular Ecology Resources 9: 1530-1535. 10.1111/j.1755-0998.2009.02710.x
Arata, J., Croxall, J.P., Huin, N., Nicholls, D., Phillips, R.A., Quintana, F., Robertson, C.J.R., Robertson, G., Trathan, P.N., Falabella, V. (2009) Albatros. In: Falabella, V., Campagna, C., Croxall, J.P. (Eds) Atlas del Mar Patagónico: Especies y espacios. Buenos Aires Wildlife Conservation Society, BirdLife International.
Bost, C.A., Thiebot, J.B., Pinaud, D., Cherel, Y., Trathan, P.N. (2009). Where do penguins go during the inter-breeding period? Using geolocation to track the winter dispersion of the macaroni penguin. Biology Letters 5: 473-476. 10.1098/rsbl.2009.0265.
Campagna, C., Fedak, M., Lewis, M., Staniland, I.J., Thompson, D., Trathan P.N., Wilson, R., Marin, M.R. Falabella, V. (2009) Pinípedos. In: Falabella, V., Campagna, C., Croxall, J.P. (Eds) Atlas del Mar Patagónico: Especies y espacios. Buenos Aires Wildlife Conservation Society, BirdLife International.
Forcada, J., Trathan, P.N. (2009). Penguin responses to climate change in the Southern Ocean. Global Change Biology 15: 1618-1630. DI 10.1111/j.1365-2486.2009.01909.x.
Ford, H., Conochie, G., Trathan, P.N., Gillett, H.J. (2009). The decline of the Bankouale palm in Djibouti: El Niño and changes in architectural fashion. Palms 52: 89-95.
Fretwell, P.T., Trathan, P.N. (2009). Penguins from space: faecal stains reveal the location of emperor penguin colonies. Global Ecology and Biogeography 18: 543-552. 10.1111/j.1466-8238.2009.00467.x.
Hart, T., Fitzcharles, E., Trathan, P.N., Coulson, T., Rogers, A.D. (2009). Testing and Improving the Accuracy of Discriminant Function Tests: A Comparison between Morphometric and Molecular Sexing in Macaroni Penguins. Waterbirds 32: 437-443.
Hill, S.L., Trathan, P.N., Agnew, D.J. (2009). The risk to fishery performance associated with spatially resolved management of Antarctic krill (Euphausia superba) harvesting. ICES Journal Marine Science 66: 2148-2154. 10.1093/icesjms/fsp172.
Quintana, F., Croxall, J.P., Gonzáles-Solis, J., Phillips, R.A., Trathan, P.N., Falabella, V. (2009). Petreles. In: Falabella, V., Campagna, C., Croxall, J.P. (Eds) Atlas del Mar Patagónico: Especies y espacios. Buenos Aires Wildlife Conservation Society, BirdLife International.
Sakamoto, K.Q., Takahashi, A., Iwata, T., Trathan, P.N. (2009). From the eye of the albatrosses: a bird-borne camera shows an association between albatrosses and a killer whale in the Southern Ocean. PLoS One 4: 7322. 10.1371/journal.pone.0007322.
Trathan, P.N., Reid, K. (2009). Exploitation of the marine ecosystem in the sub-Antarctic: historical impacts and current consequences. Papers and Proceeding of the Royal Society of Tasmania 143: 9-14.
Turner, J. et al. (2009). The Instrumental Period. In: Turner, J., Bindschadler, R., Convey, P., di Prisco, G., Fahrbach, E., Gutt, J., Hodgeson, D., Mayewski, P., Summerhays, C. (Eds) (2009). Antarctic Climate Change and the Environment. SCAR, Cambridge UK.
Wakefield, E.D., Phillips, R.A., Matthiopoulos, J., Fukuda, A., Higuchi, H., Marshall, G.J., Trathan, P.N. (2009). Wind field and sex constrain the flight speeds of central-place foraging albatrosses. Ecological Monographs 79: 663-679.

Barbraud, C., Weimerskirch, H., Bost, C-A., Forcada, J., Trathan, P.N., Ainley, D. (2008). Are king penguin populations threatened by Southern Ocean warming? Proc Natl Acad Sci U S A 105: E38; author reply E39.
Forcada, J., Trathan, P.N., Murphy, E.J. (2008). Life history buffering in Antarctic mammals and birds against changing patterns of climate and environmental variation. Global Change Biology 14: 2473-2488.
Takahashi, A., Ochi, D., Watanuki, Y., Deguchi, T., Tomohiro, O., Oka N., Afanasyev, V., Fox, J.W., Trathan, P.N. (2008). Post-breeding movement and activities of two Streaked Shearwaters in the north-western Pacific. Ornithological Science 7: 29-35.
Petit, J., Prudent, G. (Eds) (2008). Climate Change and Biodiversity in the European Union Overseas Entities. IUCN, Brussels 178pp.
Trathan, P.N., Bishop, C., Maclean, G., Brown P., Fleming, A., Collins, M.A. (2008). Linear tracks and restricted temperature ranges characterise penguin foraging pathways. Marine Ecology-Progress Series 370: 285-294.
Trathan, P.N., Forcada, J., Atkinson, R., Downie, R.H., Shears, J.R. (2008). Population assessments of gentoo penguins (Pygoscelis papua) breeding at an important Antarctic tourist site, Goudier Island, Port Lockroy, Palmer Archipelago, Antarctica. Biological Conservation 141: 3019-3028.
Yamamoto, T., Takahashi, A., Yoda, K., Katsumata, N., Watanabe, S., Sato, K., Trathan, P.N. (2008). The lunar cycle affects at-sea behaviour in a pelagic seabird, the streaked shearwater, Calonectris leucomelas. Animal Behaviour 76: 1647-1652.

Bishop, C., Trathan, P.N., MacLean, G., Brown, P. (2007) Novel GPS tags reveal fine-scale foraging movements of marine predators over extended time periods. Comparative biochemistry and physiology A - Molecular and integrative physiology 146: S86-S86 33 Suppl. S.
Cresswell, K.A., Tarling, G.A., Trathan, P.N. (2007). Weight loss during breeding is adaptive for female macaroni penguins, Eudyptes chrysolophus. Evolutionary Ecology Research 9: 1053-1076.
Hoffman, J.I., Forcada, J., Trathan, P.N., Amos, W. (2007) Female fur seals show active choice for males that are heterozygous and unrelated. Nature 445: 912-914.
Murphy, E.J., Trathan, P.N., Watkins, J.L., Reid, K, Meredith, M.P., Forcada, J., Thorpe, S.E., Johnston, N.M., Rothery, P. (2007) Climatically driven fluctuations in Southern Ocean ecosystems. Source: Proc. Roy. Soc. B 274: 3057-3067.
Murphy, E.J., Watkins, J.L., Trathan, P.N., Reid, K., Meredith, M.P., Thorpe, S.E., Johnston, N.M., Clarke, A., Tarling, G.A., Collins, M.A., Forcada, J., Shreeve, R.S., Atkinson, A., Korb, R., Whitehouse, M.J., Ward, P., Rodhouse, P.G., Enderlein, P., Hirst, A.G., Martin, A.R., Hill, S.L., Staniland, I.J., Pond, D.W., Briggs, D.R., Cunningham, N.J., Fleming, A.H. (2007) Spatial and temporal operation of the Scotia Sea ecosystem: a review of large-scale links in a krill centred food web. Phil. Tran. Roy. Soc. B 362:113-148.
Nicol, S., Croxall, J.P., Trathan, P.N., Gales, N., Murphy E.J. (2007) Paradigm misplaced? Antarctic marine ecosystems are affected by climate change as well as biological processes and harvesting. Antarct. Sci. 19 (3): 291-295
Rey, A.R., Trathan, P.N., Schiavini, A. (2007) Inter-annual variation in provisioning behaviour of southern rockhopper penguins Eudyptes chrysocome chrysocome at Staten Island, Argentina. Ibis 149: 826-835.
Rey, A.R., Trathan, P.N., Pütz, K., Schiavini, A. (2007) Effect of oceanographic conditions on the winter movements of rockhopper penguins Eudyptes chrysocome chrysocome from Staten Island, Argentina. Marine Ecology-Progress Series, 330: 285-295.
Sato, K., Watanuki, Y., Takahashi, A., Miller, P.J.O., Tanaka, H., Kawabe, R., Ponganis, P.J., Handrich, Y., Akamatsu, T., Watanabe, Y., Mitani, Y., Costa, D.P., Bost, C.A., Aoki, K., Amano, M., Trathan, P.N., Shapiro, A., Naito, Y. (2007) Stroke frequency, but not swimming speed, is related to body size in free-ranging seabirds, pinnipeds and cetaceans. Proc. Roy. Soc. B 274: 471-477.
Trathan, P.N., Forcada J., Murphy E.J. (2007) Environmental Forcing and Southern Ocean marine predator populations: effects of climate change and variability. Phil. Tran. Roy. Soc. 362: 2351-2365.

Forcada, J., Trathan, P.N., Reid, K., Murphy, E.J., Croxall, J.P. (2006) Contrasting population changes in sympatric penguin species in association with climate warming. Global Change Biology 12: 411-423.
Green, C.J., Trathan, P.N., Preston, M. (2006) A new automated logging gateway to study the demographics of Macaroni penguins (Eudyptes chrysolophus) at Bird Island, South Georgia: Testing the reliability of the system using radio telemetry. Polar Biology 29: 1003-1010.
Hill, S.L., Murphy, E.J., Reid, K., Trathan, P.N., Constable, A.J. (2006) Modelling Southern Ocean ecosystems: krill, the food-web, and the impacts of harvesting. Biological Reviews 81: 581-608.
Hoffman, J.I., Trathan, P.N., Amos, W. (2006) Genetic tagging reveals extreme site fidelity in territorial male Antarctic fur seals Arctocephalus gazelle. Molecular Ecology 15: 3841-3847.
Leaper, R., Cooke, J., Trathan, P.N., Reid, K., Rowntree, V., Payne, R. (2006) Global climate drives southern right whale (Eubalaena australis) population dynamics. Biology Letters 2: 289-292, doi:10.1098/rsbl.2005.0431.
Poncet S., Robertson G., Phillips R.A., Lawton K., Phalan B., Trathan P.N., Croxall J.P. (2006) Status and distribution of wandering, black-browed and grey-headed albatrosses breeding at South Georgia. Polar Biology 29: 772-781.
Reid, K., Murphy, E.J., Croxall, J.P., Trathan, P.N. (2006) Population dynamics of Antarctic krill Euphausia superba at South Georgia - sampling with predators provides new insights In: Boyd, I.L., Wanless, S., Camphuysen, C.J. (Eds.), Top Predators in Marine Ecosystems, Cambridge University Press.
Staniland, I.J., Trathan, P.N., Martin, A.R. (2006) Consequences of prey distribution for the foraging behaviour of top predators In: Boyd, I.L., Wanless, S., Camphuysen, C.J. (Eds.), Top Predators in Marine Ecosystems, Cambridge University Press.
Trathan, P.N., Green, C., Tanton, J., Peat, H., Poncet, J., Morton A. (2006) Foraging dynamics of macaroni penguins Eudyptes chryolophus at South Georgia during brood-guard. Marine Ecology-Progress Series 323: 239-251.
Trathan, P.N., Murphy, E. J., Forcada, J., Croxall, J. P., Reid, K., Thorpe, S.E. (2006) Physical forcing in the southwest Atlantic: ecosystem control. In: Boyd, I.L., Wanless, S., Camphuysen, C.J. (Eds.), Top Predators in Marine Ecosystems, Cambridge University Press.
Warren N.L., Trathan P.N., Forcada J., Fleming A., Jessopp M.J. (2006) Distribution of post-weaning Antarctic fur seal Arctocephalus gazelle pups at South Georgia. Polar Biology, 29: 179-188.

Forcada, J., Trathan, P.N., Reid, K., Murphy, E.J. (2005) The effects of global climate variability in pup production of Antarctic fur seals. Ecology, 86: 2408-2417.
Meredith, M.P., Brandon, M.A., Murphy, E.J., Trathan, P.N., Thorpe, S.E., Bone, D.G., Chernyshkov, P.P., Sushin, V.A. (2005) Variability in hydrographic conditions to the east and northwest of South Georgia, 1996-2001. J. Mar. Syst., 53: 143- 167.

Brandon, M.A., Naganobu, M., Demer, D.A., Chernyshkov, P., Trathan, P.N., Thorpe, S.E., Kameda, T., Berezhinskiy, O.A., Hawker, E.J., Grant, S. (2004) Physical oceanography in the Scotia Sea during the CCAMLR 2000 survey, austral summer 2000. Deep-Sea Res. Part II, 51: 1301-1321.
Croxall, J.P., Trathan, P.N. (2004) The Southern Ocean: a model system for conserving resources? (In Glover, L.K. & Earle, S.A., eds. Defying Ocean's end: an agenda for action. Washington D.C., Island Press, pp 69-87.
Hewitt, R.P., Watkins, J.L., Naganobu, M., Sushin, V., Brierley, A.S., Demer, D., Kasatkina, S., Takao, Y., Goss, C., Malyshko, A., Brandon, M.A., Kawaguchi, S., Siegel, V., Trathan, P.N., Emery, J., Everson, I., Miller, D.G. (2004) Biomass of Antarctic krill in the Scotia Sea in January/February 2000 and its use in revising an estimate of precautionary yield. Deep-Sea. Res. Part II. 51: 1215-1236.
Hewitt, R.P., Watters, G., Trathan, P.N., Croxall, J.P., Goebel, M.E., Ramm, D., Reid, K., Trivelpiece, W.Z., Watkins, J.L. (2004) Options for allocating the precautionary catch limit of krill among small-scale management units in the Scotia Sea. CCAMLR Sci. 11: 81-97.
Jessopp, M.J., Forcada, J., Reid, K., Trathan, P.N., Murphy, E.J. (2004) Winter dispersal of leopard seals (Hydrurga leptonyx) environmental factors influencing demographics and seasonal abundance. J. Zool., 263: 251-258.
Murphy, E.J., Watkins, J.L., Meredith, M.P., Ward, P., Trathan P.N., Thorpe, S.E. (2004) Southern Antarctic Circumpolar Current Front to the northeast of South Georgia: Horizontal advection of krill and its role in the ecosystem. J. Geophys. Res., 109: art. no. C01029.
Nevitt, G., Reid, K., Trathan, P.N. (2004) Testing olfactory foraging strategies in an Antarctic seabird Assemblage. J. Exp. Biol. 207: 3537-3544.
Tanton, J.L., Reid, K., Croxall, J.P., Trathan, P.N. (2004) Winter distribution and behaviour of gentoo penguins Pygoscelis papua at South Georgia. Polar Biol., 27: 299-303.
Takahashi, A., Sato, K., Naito, Y., Dunn, M.J., Trathan, P.N., Croxall, J.P. (2004) Penguin-mounted cameras glimpse underwater group behaviour. Proc. Roy. Soc. Lond. Ser. B., 271: S281-S282.
Trathan, P.N. (2004) Image analysis of color aerial photography to estimate penguin population size. Wildlife Soc. Bull., 32: 332-343.
Trathan, P.N., Croxall, J.P. (2004) Marine predators at South Georgia: an overview of recent bio-logging studies. Memoirs of National Institute of Polar Research. 58: 118-132.
Waluda, C.M., Trathan, P.N., Rodhouse, P.G. (2004) Synchronicity in southern hemisphere squid stocks and the influence of the Southern Oscillation and Trans Polar Index. Fish. Oceanog., 13: 255-266.
Xavier, J.C., Trathan, P.N., Croxall, J.P., Wood, A.G., Podesta, G., Rodhouse, P.G. (2004) Foraging ecology and interactions with fisheries of wandering albatrosses (Diomedea exulans) breeding at South Georgia. Fish. Oceanog., 13: 324-344.

Croxall, J.P., Trathan, P.N., Murphy, E.J. (2003) Adélie penguins and environmental change – Response. Science, 300: 429-430.
Takahashi, A., Dunn, M.J., Trathan, P.N., Sato, K., Naito, Y., Croxall, J.P. (2003) Foraging strategies of chinstrap penguins at Signy Island, Antarctica: importance of benthic feeding on Antarctic krill. Mar. Ecol.-Prog. Ser., 250: 279-289.
Trathan, P.N., Brierley, A.S., Brandon, M.A., Bone, D.G., Goss, C., Grant, S.A., Murphy, E.J., Watkins, J.L. (2003) Oceanographic variability and changes in Antarctic krill (Euphausia superba) abundance at South Georgia. Fish. Oceanog.,12: 569-583.
Xavier, J.C., Croxall, J.P., Trathan, P.N., Rodhouse, P.G. (2003) Inter-annual variation in the cephalopod component of the diet of the wandering albatross, Diomedea exulans breeding at Bird Island, South Georgia. Mar. Biol., 142: 611-622.
Xavier, J.C., Croxall, J.P., Trathan, P.N., Wood, A.G. (2003) Feeding strategies and diets of breeding grey-headed and wandering albatrosses at South Georgia. Mar. Biol., 143: 221-232.

Croxall, J.P., Trathan, P.N., Murphy, E.J. (2002) Environmental change and Antarctic seabird populations. Science, 297: 1510-1514.
Lynnes, A.S., Reid, K., Croxall, J.P., Trathan, P.N. (2002) Conflict or co-existence? Foraging distribution and competition for prey between Adélie and chinstrap penguins. Mar. Biol., 141: 1165-1174.
Trathan, P.N., Murphy, E.J. (2002) Sea surface temperature anomalies near South Georgia: Relationships with the pacific El Niño regions. J. Geophys. Res., 108: art. no. 8075.
Waluda, C.M., Trathan, P.N., Elvidge, C.D., Hobson, V.R., Rodhouse, P.G. (2002) Throwing light on straddling stocks of Illex argentinus: assessing fishing intensity with satellite imagery. Can. J. Fish. Aquat. Sci., 59: 592-596.

Rodhouse, P. G. K., Elvidge, C. and Trathan, P. N. (2001) Remote sensing of the global light-fishing fleet: an analysis of interactions with oceanography, other fisheries and predators. Advances in Marine Biology, 39: 261-303
Trathan, P. N., Watkins, J. L., Murray. A. W. A., Hewitt, R. P., Naganobu, M., Sushin, V., Brierley, A. S., Demer, D., Everson, I., Goss, C., Hedley, S., Katsakina, S., Kawaguchi, S., Kim, S., Pauly, T., Priddle, J., Reid, K. and Ward, P. (2001). The CCAMLR-2000 krill synoptic survey; a description of the rationale and design. CCAMLR Science 8: 1-24
Waluda, C. M., Rodhouse, P. G., Trathan, P. N. and Pierce, G. J. (2001) Remotely sensed mesoscale oceanography and distribution of Illex argentinus: in the South Atlantic. Fish. Oceanog. 10: 207-216
Waluda, C. M., Rodhouse, P. G., Podestá, G. P., Trathan, P. N. and Pierce, G. J. (2001) Surface oceanography of the inferred hatching grounds of Illex argentinus (Cephalopoda: Ommastrephidae) and influences on recruitment variability. Mar. Biol. 139: 671-679

Brandon, M. A., Murphy, E. J., Trathan, P. N. and Bone, D. G. (2000). Physical oceanographic conditions to the northwest of the sub-Antarctic Island of South Georgia. J. Geophys. Res. 105: 23983-23996
Trathan, P. N., Brandon, M. A., Murphy, E. J. and Thorpe, S. E. (2000). Transport and structure within the Antarctic Circumpolar Current to the north of South Georgia. Geophys. Res. Lett., 27: 1727-1730

Brandon, M. A., Murphy, E. J., Whitehouse, M. J., Trathan, P. N., Murray. A. W. A, Bone, D. G. and Priddle, J. (1999) The shelf break front to the east of the sub-Antarctic island of South Georgia. Cont. Shelf Res. 19: 799-819
Trathan, P. N., Kalinowski, J., Everson, I. (1999) Status of the Polish FIBEX acoustic data from the West Atlantic. CCAMLR Science 6: 125-132
Walluda C. M., Trathan, P. N. and Rodhouse, P. G. (1999) Influence of oceanographic variability on recruitment in the Illex argentinus (Cephalopoda: Ommastrephidae) fishery in the South Atlantic. Mar. Ecol. Prog. Ser. 183: 159-167
Xavier, J. C., Rodhouse, P. G. K., Trathan, P. N. and Wood, A. G. (1999) A Geographical Information System (GIS) Atlas of squid distribution in the Southern Ocean. Antarct. Sci. 11: 61-62


Murphy, E. J., Watkins, J. L., Reid, K., Trathan, P. N., Everson, I., Croxall, J. P., Priddle, J., Brandon, M. A., Brierley, A. S. and Hofmann, E. H. (1998) Interannual variability of the South Georgia marine ecosystem: biological and physical sources of variation in the abundance of krill. Fish. Oceanog. 7: 381-390
North. A. W., White, M. G and Trathan, P. N. (1998) Interannual variability in the early growth rate and size of the Antarctic fish Gobionotothen gibberifrons (Lonnberg) Antarctic. Sci. 10: 416-422
Prince, P. A., Croxall, J. P., Trathan, P. N. and Wood, A. G. (1998) The pelagic distribution of South Georgia albatrosses and their relationships with fisheries, in Albatross Biology and Conserva¬tion, edited by G. Robertson, and Gales, R., Surrey Beatty and Sons, Chipping Norton, Australia, 137-167
Trathan, P. N., Everson, I., Murphy, E. J. and Parkes, G. (1998) Analysis of haul data from the South Georgia krill fishery. CCAMLR Science 5: 9-30
Trathan, P. N., Murphy, E. J., Croxall, J. P. and Everson, I. (1998) Use of at-sea distribution data to derive potential foraging ranges of macaroni penguins during the breeding season. Mar. Ecol. Prog. Ser. 169: 263-275

Gonzalez, A. F., Trathan, P. N., Yao, C. and Rodhouse, P. G. K. (1997) Interactions between oceanography, ecology and fishery biology of the ommastrephid squid Martialia hyadesi in the South Atlantic. Mar. Ecol. Prog. Ser. 152: 205-215
Murphy, E. J., Trathan, P. N., Everson, I., Daunt, F. H. J. and Parks, G. (1997) Krill fishing distribution around South Georgia. CCAMLR Science 4: 1-18
Trathan, P. N., Brandon, M. A. and Murphy, E. J. (1997) Characterisation of the Antarctic Polar Frontal Zone to the north of South Georgia in summer 1994. J. Geophys. Res. 102: C5 10483-10497

Hill, H. J., Trathan, P. N., Croxall, J. P. and Watkins, J. L. (1996) A comparison of Antarctic krill, Euphausia superba, caught by nets and by macaroni penguins, Eudyptes chrysolophus: evidence for selection? Mar. Ecol. Prog. Ser. 140: 1-11
Reid, K., Trathan, P. N. Croxall, J. P. and Hill, H. J. (1996) Krill caught by predators and nets: differences between species and techniques. Mar. Ecol. Prog. Ser. 140: 13-20
Rodhouse, P. G. K., Prince, P. A., Trathan, P. N., Hatfield, E. M. C., Watkins, J. L., Bone, D. G., Murphy, E. J. and White, M. G. (1996) Cephalopod community associated with mesoscale oceanographic features at the Antarctic Polar Front exploited by satellite tagged predators. Mar. Ecol. Prog. Ser. 136: 37-50
Trathan, P. N., Croxall, J. P. and Murphy, E. J. (1996) Dynamics of Antarctic penguin populations in relation to the inter-annual variability in sea-ice distribution. Polar Biology 16: 321-330
Ward, P., Shreeve, R. S., Cripps, G. C. and Trathan, P. N. (1996) Mesoscale distribution and population dynamics of Rhincalanus gigas and Calanus simillimus in the Antarctic Polar Open Ocean and Polar Frontal Zone during summer. Mar. Ecol. Prog. Ser. 140 21-32
Whitehouse, M. J., Priddle, J., Trathan, P. N. and Brandon, M. A. (1996) Substantial open-ocean phytoplankton blooms to the north of South Georgia, South Atlantic, during summer 1994. Mar. Ecol. Prog. Ser. 140: 187-197


Trathan, P. N. and Everson, I. (1995) Status of the FIBEX acoustic data from the West Atlantic. CCAMLR Science 1: 35-48
Trathan, P. N., Everson, I., Miller, D. G. M., Watkins, J. L. and Murphy, E. J. (1995) Krill biomass in the Atlantic. Nature 367: 201-202


Thorley, M. R. and Trathan, P. N. (1994) The BIOMASS dataset documentation. British Antarctic Survey. ISBN 0 85665 159 1

Hunt, G. L., Croxall, J. P. and Trathan, P. N. (1993) Marine Ornithology in the southern Drake Passage and Bransfield Strait during the BIOMASS programme in El-Sayed SZ (Ed.) Southern Ocean Ecology: The BIOMASS Perspective, Cambridge University Press.
Priddle, J., Thorley, M. R. and Trathan, P. N. (1993) The potential contributions of the BIOMASS programme to global change research, especially to the JGOFS core project of IGBP in El-Sayed SZ (Ed.) Southern Ocean Ecology: The BIOMASS Perspective, Cambridge University Press.
Thorley, M. R. and Trathan, P. N. (1993) The history of the BIOMASS Data Centre and lessons learned during its lifetime in El-Sayed SZ (Ed.) Southern Ocean Ecology: The BIOMASS Perspective, Cambridge University Press.
Trathan, P. N., Agnew, D., Miller, D. G. M., Watkins, J. L., Everson, I., Thorley, M. R., Murphy, E. J., Murray, A. W. A. and Goss, C. (1993a) Krill Biomass in Area 48 and Area 58: Recalculation of FIBEX Data. SC-CAMLR-SSP/9 WG-KRILL-92/20: 157-181
Trathan, P. N., Murphy, E. J., Symon, C. J. and Rodhouse, P.G. K. (1993b) Ecological and oceanographic relationships in the Southern Ocean. GIS Europe 2: 34-36
Trathan, P. N., Priddle, J., Watkins, J. L., Miller, D. G. M. and Murray, A. W. A. (1993c) Spatial variability of Antarctic krill in relation to mesoscale hydrography. Mar. Ecol. Prog. Ser. 98: 61-71

Pre 1993
Gliddon, C. J. and Trathan, P. N. (1985) Interactions between white clover and perennial ryegrass in an old permanent pasture in Haeck, J. and Woldendorp, J.W. (eds.) Phenotypic and genotypic variation in plant populations.
Trathan, P. N. (1983) Clonal interactions of Lolium perenne and Trifolium repens in an old permanent pasture. PhD Thesis, University of Wales.